Sudanese protest leaders on Wednesday turned down an offer by the ruling military council for talks and demanded justice for a crackdown that doctors said has left 108 people dead.
Security forces moved in to brutally disperse a protest sit-in on Monday.
The Rapid Support Forces, paramilitaries said by rights groups to have their origins in the Janjaweed militias accused of abuses during the 16-year-old conflict in Darfur, are thought to have been largely behind the crackdown.
The Central Committee for Sudanese Doctors close to the protest movement said on Wednesday that at least 108 people had been killed in the crackdown, including 40 whose bodies were recovered from the Nile, and more than 500 wounded.
The Sudan Doctors' Committee said Wednesday the bodies were pulled out a day earlier, and that they were taken by the paramilitary Rapid Support Forces to an unknown location. The sit-in was broken up by the military on Monday.
One activist, Amal al-Zein, said the number could be even higher. She said dozens of bodies had been pulled from the Nile in different places near the sit-in area and were taken to a hospital morgue.
People 'beaten and thrown in Nile'
"Some bodies have wounds from bullets, others seemed to have beaten and thrown in the Nile," she said.
The announcement came as Sudanese protest leaders dismissed a call for talks with the ruling generals, saying the military cannot be serious about negotiations while troops keep shooting and killing protesters.
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"The Sudanese people are not open for talks, the Sudanese people are not open to this TMC (Transitional Military Council) that kills people and we need justice and accountability before talks about any political process," Amjad Farid, a spokesman for the Sudanese Professionals Association, which spearheaded protests that led to the ouster of president Omar al-Bashir, told AFP.
Farid said the SPA and umbrella protest group the Alliance for Freedom and Change would "continue using all non-violent tools and civil disobedience in resisting the TMC".
For its part, the United Nations has responded to this latest spell of violence by deciding to remove some of its staff from the country on Wednesday: "We are temporarily relocating non-programme-critical UN staff, while all UN operations continue in Sudan," said UN spokesperson Eri Kaneko.
Earlier the same day, the head of the military council, Gen. Abdel-Fattah Burhan, said the generals were ready to resume negotiations and that there would be "no restrictions" in talks with the leaders behind the months-long street protests.
"We open our hands to negotiations with all parties ... for the interest of the nation," Burhan said, adding that those responsible for the violent beak-up of the demonstrators' sit-in in the capital, Khartoum, would be held accountable.
The motives for Burhan's about-face if sincere were not immediately clear. Burhan had earlier cut the negotiations and canceled all agreed-on points between the military and the Forces for Declaration of Freedom and Change, an alliance which represents the protesters.
Mohammed Yousef al-Mustafa, a spokesman for the Sudanese Professionals Association, which is also part of the alliance and which is leading the demonstrations, said the protesters "totally reject" Burhan's call.
"This call is not serious," al-Mustafa told The Associated Press. "Burhan and those under him have killed the Sudanese and are still doing it. Their vehicles patrol the streets, firing at people."
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"We will continue in our protests, resistance, strike and total civil disobedience," he added.
Meanwhile, the death toll in the military's crackdown reached 100 on Wednesday, organisers said. The mounting casualties are the latest challenge to the protest movement, which now aims to show it can keep up pressure in the streets after its central rallying point the weeks-long sit-in outside the military headquarters in Khartoum was wiped out on Monday.
Ongoing sit-ins after Bashir's ouster
In April, the movement succeeded in forcing the military to remove Sudan's longtime strongman, Omar al-Bashir. It then kept its sit-in going, demanding that the generals who took power hand over authority to civilians.
The last previously reported death toll stood at 40 but the doctors committee said security forces killed at least 10 people on Wednesday in Khartoum and its twin city of Omdurman. That came after another 10 people were killed on Tuesday, including five in the White Nile state, three in Omdurman and two in Khartoum's Bahri neighborhood. That total 100 includes the 40 bodies pulled from the river.
The doctors' committee, the medical arm of the Sudanese Professionals Association, also said that at least 326 people have been wounded in clashes in the past two days and that it feared the final death toll would be much higher.
Activists Mohammed Najib and Hashim al-Sudani said there were street battles late Tuesday and early Wednesday in Khartoum's Bahri and Buri districts between protesters and security forces, mainly from the paramilitary Rapid Support Forces, or RSF.
"In Buri, there were lots of shootings and tear gas," al-Sudani said. "They tried to force people into narrow streets" to beat them.
The opposition Congress Party reported more protests on Wednesday, posting footage showing hundreds of people marching in Omdurman.
On Monday, members of the RSF, which human rights groups say carried out rapes, torture and killings of civilians in Sudan's Darfur region, and other troops waded into the protest camp outside the military's headquarters in Khartoum, opening fire and burning down tents. Other troops crushed two smaller sit-ins organised by the protesters elsewhere.
Weeks of negotiations
After al-Bashir's ouster, for weeks, the military and protest leaders negotiated over the makeup of a transitional council meant to run the country for three years before elections. Protesters demand that civilians dominate the council, but the generals resisted.
The crackdown put an end to the relative peace that surrounded the talks and signaled the military had lost patience with activists' demands, putting the two sides on the path of a potentially longer confrontation with increasing violence.
Burhan's call for negotiations marked a significant change from his televised speech only a day earlier, when he blamed protest leaders for the volatile situation, accusing them of drawing out negotiations and trying to exclude some "political and security forces" from taking part in any transitional government.
He also announced Tuesday that the military would unilaterally form an interim government and hold elections sooner, within seven to nine months, under international supervision. Protesters had also rejected that because it would put the military in charge of running the election.
Since Monday, tensions have remained high in Khartoum and elsewhere as demonstrators staged scattered rallies and blocked main roads with barricades. Security forces have been seen in online videos, patrolling Khartoum districts.
Madani Abbas Madani, a leading activist, said the protesters would continue an open-ended civil disobedience campaign until the overthrow of the ruling military council.
"What happened on Monday was a systematic and planned attempt to impose repression on the Sudanese people," he said.
In a joint statement, the US, Norway and Britain have condemned the security forces' crackdown. The statement, released late Tuesday, said Sudan's military council "put the transition process and peace in Sudan in jeopardy" by ordering attacks on protesters.
(FRANCE 24 with AP and AFP)
