Devin Toner will start in Ireland's Six Nations clash with England on Sunday.
The veteran second row replaces Iain Henderson, who has withdrawn from the squad due to family reasons, with his wife giving birth to a baby boy this week.
Toner's last start for Ireland in the Six Nations was at home to Italy last year, and it was a shock that the long-serving lock missed out on World Cup selection in the autumn.
Toner's 70th cap follows appearances off the bench against Scotland and Wales in the opening two championship fixtures.
His place on the bench goes to Connacht's Ultan Dillane, who last played in the Six Nations in the victory over France last year.
"Faz (Andy Farrell) was speaking with Iain during the week, they spoke a bit last night and it became clearer this morning, and we planned during the week in the case that it happened," explained Ireland scrum coach John Fogarty today.
"It would have been an ask for him to come back in after missing two sessions. We're delighted for him, a little baby boy.
"And the boys here have had good prep this week. We're delighted for Dev, he missed the World Cup and what a game to come back into as a starter.
"They're primed and ready and will need to be, but we're looking forward to it."
Toner's place on the bench goes to Connacht's Ultan Dillane, who last played in the Six Nations in the victory over France last year.
Skipper Johnny Sexton is delighted to see Toner back from the start as Ireland look to clinch the Triple Crown.
"It's brilliant for him," said Sexton.
"Dev had a big setback at a bad time for him last year but he got some reception against Scotland when he came on, he got a bit of stick for it!
"England have made a few changes, a few unexpected things, but we'll catch up on those this afternoon and this evening. England have so much depth, and they're very lucky to have it.
"We're going to be preparing for the best England that we've seen in this championship. But you can't have more motivation than going to Twickenham looking for the Triple Crown.
"We have to make up a 50-point margin from when we last played them and it's been a good week figuring that out."
IRELAND: Larmour, Conway, Henshaw, Aki, Stockdale, Sexton, Murray; Healy, Herring, Furlong, Ryan, Toner, O'Mahony, van der Flier, Stander. Replacements: Kelleher, Kilcoyne, Porter, Toner, Doris, Cooney, Byrne, Earls.
