The last Sumatran rhinoceros in Malaysia died today - leaving the smallest species of rhino now surviving in tiny numbers in Indonesia.
The 25-year-old animal, named Iman, began to suffer "significant pain" after falling unwell to cancer.
She had been cared for in a wildlife reserve in the state of Sabah on the island of Borneo since her capture in 2014.
"Iman's death came rather sooner than we had expected, but we knew that she was starting to suffer significant pain," said Augustine Tuuga, director of the Sabah Wildlife Department.
The Sumatran rhino was declared extinct in the wild in Malaysia in 2015.
The country's last male Sumatran rhino died in May this year.
There are now thought to be less than 80 Sumatran rhino in the wild in Indonesia.
Conservationists believe they mainly live on the Indonesian island of Sumatra and on the Indonesian side of Borneo.
But habitat loss and poaching endangered them years ago.
They rarely breed as a result, conservation group International Rhino Foundation says.
Malaysia has, since 2011, tried to breed the species in captivity through in-vitro fertilization, but without success.
When they were declared extinct in the wild in the nation in 2015, scientists predicted a grim outlook.
Rasmus Gren Havmøller, from the Center for Macroecology, Evolution and Climate, said at the time: "It is vital for the survival of the species that all remaining Sumatran rhinos are viewed as a metapopulation, meaning that all are managed in a single program across national and international borders in order to maximize overall birth rate.
"This includes the individuals currently held in captivity."
