Owen Miller was a rank outsider heading into the T20 1500m final, but upset the odds to fend off stiff competition from the Russian and American favourites to claim a shock gold medal - ParalympicsGB's first of day 10.
The 29-year-old crossed the line in 3:54.57, as he ran to victory at a very wet Olympic Stadium in Tokyo.
In the end he streaked away from world and European champion Alexandr Rabotnitskii of the Russian Paralympic Committee, who was expected to tussle with reigning Paralympic champion Michael Brannigan of Team USA for gold.
Brannigan finished the race with a season's best time of 3:58.43, but that wasn't even enough for a medal in what proved to be a very fast race.
Miller and Rabotnitskii broke away from the group as they came around the last bend and the Russian had the inside line, but the Scot came bursting around the outside and pulled away as the world champion could not match his pace.
Miller finished more than a second ahead of Rabotnitskii, while Ndiaga Dieng of Italy claimed bronze ahead of Brannigan.
In the women's race there was also success for Britain, as Hannah Taunton ran a personal best 4:35.34 to add a bronze medal to Millar's gold.
While the reigning champion could not repeat the feat in the men's race, it was not the same for the women as Barbara Bieganowska-Zajac of Poland won the race for the third Games in a row.
Another gold for ParalympicsGB came courtesy of Emma Wiggs in the women's VL2 canoe.
And, in the T64 high jump, Jonathan Broom-Edwards' 2.10m season's best was a silver medal-winning score five years ago, but it was enough to take the crown in Tokyo as the 33-year-old claimed his first Paralympic title and took Britain's gold medal total to 37.
"I am so emotional. To just get it right at the right time, I am so relieved, elated, excited and crying my eyes out," he said.
"I've been striving for that gold for years. They were horrible conditions so I tried to keep my cool and get it right when it counted. And I am so proud of myself.
"(Silver medallist) Praveen (Kumar) pulled out a personal best and fair play to him, that put me on the back foot.
"I had to rise above that and I am so proud that I did. 2.10m was a bit of an ugly jump but it was a season best so I am delighted."
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